The soils of a rainforest affect the diversity of the forest.
Although nearly 70 percent of tropical rainforest exists on
poor acidic soils, it retains its fertility in a large part thanks to
nutrient recycling and other processes. However, in some areas, soils
are so poor that only a limited number of tree species can grow (though
these forests are still highly diverse by temperate standards). One
example is the so-called "white-sand" or "blackwater"
forests that grow on rocky, sandy soils. Some of these forests grow
on nothing but rocks and the roots of other trees. Trees that grow under
these conditions tend to be species with tannins in their leaves, which
in turn, turn local rivers into "blackwater" rivers. The bitter
tannins in their leaves limit insect populations, thus reducing
the number of animals the forest can support (insects serve as a major
food source for larger animals in most rainforests). These "blackwater"
forests are self-perpetuating, since the "blackwater" rivers
that result from the decay of their leaves only make the soils more
acidic and prevent other tree species from growing
on the already nutrient-lacking soils.
Forest tree diversity, and hence total diversity, may also be reduced
in forests with soggy soils like those of the igapò or "swamp
forest." The limited number of tree species like Cecropia and palms
that can tolerate these wet soil conditions means that these few trees
species tend to dominate these areas. Subsequently only the animals
that feed on their fruits, leaves, and seeds are abundant in these areas.
High-diversity forests are often found on nutrient rich—sometimes volcanic—
soils that are well-drained. These forests are frequently found in areas
protected from major disturbances like strong wind and regular flooding.
Rainforest on Barro Colorado Island, Panama. (Photo by R. Butler)