Tropical rainforests
across the world are quite diverse, but share several defining characteristics
including climate, precipitation, canopy structure, complex symbiotic
relationships, and diversity of species. Every rainforest does not necessarily
conform to these characteristics and most tropical rainforests do not
have clear boundaries, but may blend with adjoining mangrove forest,
moist forest, montane forest, or tropical deciduous forest.
GEOGRAPHY AND CLIMATE
Tropical rainforests lie in the "tropics," between the Tropic
of Capricorn and Tropic of Cancer. In this region sunlight strikes Earth
at roughly a 90-degree angle resulting in intense solar energy (solar
energy diminishes as you move farther north or south).
This intensity is due to the consistent day length on
the equator: 12 hours a day, 365 days per year (regions away from the
equator have days of varying length). This consistent sunlight provides
the essential energy necessary to power the forest via photosynthesis.
Because of the ample solar
energy, tropical rainforests are usually warm year round with temperatures
from about 72-93F (22-34C), although forests at higher elevations, especially
cloud forests, may be significantly cooler. The temperature may fluctuate
during the year, but in some equatorial forests the average may vary as little as 0.5F (0.3C) throughout the year. Temperatures
are generally moderated by cloud cover and high humidity.
PRECIPITATION
An important characteristic of rainforests is apparent in their name.
Rainforests lie in the intertropical convergence zone where intense
solar energy produces a convection zone of rising air that loses its
moisture through frequent rainstorms. Rainforests are subject to heavy
rainfall, at least 80 inches (2,000 mm), and in some areas over 430
inches (10,920 mm) of rain each year.
In equatorial regions, rainfall
may be year round without apparent "wet" or "dry"
seasons, although many forests do have seasonal rains. Even in seasonal
forests, the period between rains is usually not long enough for the
leaf litter to dry out completely. During the parts of the year when
less rain falls, the constant cloud cover is enough to keep the air
moist and prevent plants from drying out. Some neotropical rainforests rarely
go a month during the year without at least 6" of rain. The stable
climate, with evenly spread rainfall and warmth, allows most rainforest
trees to be evergreen—keeping their leaves all year and
never dropping all their leaves in any one season.
Forests further from the equator, like those of Thailand, Sri Lanka,
and Central America, where rainy seasons are more pronounced, can only
be considered "semi-evergreen" since some species of trees
may shed all of their leaves at the beginning of the dry season. Annual
rainfall is spread evenly enough to allow heavy growth of broad-leafed
evergreen trees, or at least semi-evergreen trees.
The moisture of the rainforest from rainfall, constant cloud cover,
and transpiration (water loss through leaves), creates intense local
humidity. Each canopy tree transpires some 200 gallons (760 liters)
of water annually, translating to roughly 20,000 gallons (76,000 L)
of water transpired into the atmosphere for every acre of canopy trees.
Large rainforests (and their humidity) contribute to the formation of
rain clouds, and generate as much as 75 percent of their own
rain. The Amazon rainforest is responsible for creating as much as 50
percent of its own precipitation.
The opening quotation is found in One River (New York: Touchstone, 1996) by Wade Davis.
Timme, S. ("Neotropical Plants and Ecology," Rainforest Workshop Packet 1994) pegs transpiration of individual canopy trees at 200 gallons of water per year - translating to 20,000 gallons transpired per acre of forest.
The Woods Hole Research Center concluded in its RisQue98 (Risco de Queimada, or "Risk of Burning" in Amazonia - 1998) that as much as 50% of the Amazon rainforest was at risk of burning. The report was picked up by the popular media in Lewan, Todd. "Fears of a fiery Amazon nightmare-7-year study has implications for the global warming debate," Associated Press, 12/7/97.
Two-thirds of the world's rainforests are fragmented according to M. McKloskey in "Note on the Fragmentation of Primary Rainforest," Ambio 22 (4), June: 250-51 1993.
Paine, R. T. ("Food web complexity and species diversity" American Naturalist 100: 65-75, 1966) put forth the notion of a keystone species. Further discussion of the term can be found in Power, M. E., et al., "Challenges in the quest for keystones," BioScience 46: 609-620, 1996; and Khanina, L., "Determining keystone species," Conservation Ecology 2(2), 1998.